Inhabit rocky pools and nearby riffles of headwaters and creeks (Ref. 5723); also found in streams (Ref. 10294). Often found in very shallow water less than 4 m deep (Ref. 5723). Juveniles feed on midge larvae and microcrustaceans; adults on midge larvae, other immature aquatic insects (mayflies, caddisflies, stoneflies), isopods and amphipods (Ref. 10294). Eggs are found clustered on underside of stone and guarded by males (Ref. 7043).