Inhabits rocky riffles of cold streams, rocky areas of lakes (commonly at 90-106 m depth), springs and their effluents (Ref. 5723). Moves into shallow water to spawn (Ref. 27547). In some areas, they are common in brackish water (Ref. 27843), presumably moving to and from fresh water, at least for spawning (Ref. 27547). Feeds mostly on aquatic insect larvae and nymphs but also on crustaceans, small fishes, and plant materials (Ref. 1998). May be used as bait, but this seems rare (Ref. 27547).