Inhabit shallow gravel riffles, sometimes rocky runs and pools, of headwaters, creeks and small rivers (Ref. 5723); also found in streams (Ref. 10294). Adults feed on midge and blackfly larvae, mayfly nymphs, isopods, amphipods, and caddisfly larvae (Ref. 10294). Distinct pairing during breeding (Ref. 36980). Eggs are found buried in the substrate (Ref. 7043).