Occur in quiet, vegetated lakes, headwaters, creeks and small rivers (Ref. 5723); also found in streams and springs to ponds (Ref. 10294). Usually found over mud and sand (Ref. 5723, 10294). Adults feed on midge larvae, isopods, and microcrustaceans, particularly copepods (Ref. 10294). Eggs are found attached to the substrate unguarded (Ref. 7043).