spines: 0. Deep body (Ref. 43281). Belly with 28 to 30 scutes. Head large; a median notch in upper jaw which distinguishes it from other similar clupeids, except Hilsa kelee. gill rakers fine and numerous, 204 to 316 on lower part of arch (increasing with size of fish); with mucosal buds and not asperities on upper edges of rakers. Caudal fin moderate. A dark spot behind gill opening; a series of spots along flank (Ref. 188).