HomeFish encyclopediaRay-finned fishesPerch-like fishGobiesGranular pugolovka Granular pugolovka (Benthophilus granulosus) Scientific name: Benthophilus granulosus Kessler, 1877 The Granular pugolovka (Benthophilus granulosus) is a freshwater fish of the family Gobiidae. Native range: Former USSR and Asia, Caspian Sea and the lower Volga. BiologyDorsal spines (total): 4; Dorsal soft rays (total): 7-9; Anal spines: 1; Anal soft rays: 6 - 8. Head flat, slightly depressed. Short and flat leathery barbel in chin, tubercles absent. Body color variable depending on habitat. Distinguished further from its congeners by the following characters: body covered with numerous, densely-set thorny granules; 3 dark blotches along midline of back and flank; head and back with dark dots, sometimes on dorsal, pectoral and caudal.Source: FishBase, licensed CC BY-NC 4.0. Species factsScientific nameBenthophilus granulosusFamilyGobiidae (Gobies)OrderPerciformes (Perch-like fish)ClassActinopterygii (Ray-finned fishes)Maximum length5.6EnvironmentMarine; freshwater; brackish; demersalClimateTemperateDistributionFormer USSR and Asia: Caspian Sea and the lower Volga.Conservation status (IUCN)Least Concern (LC)Threat to humansHarmlessUsesFisheries:ResilienceHigh, minimum population doubling time less than 15 months (Preliminary K or Fecundity.) SourcesTaxonomy, distribution and biology data: FishBase (Froese, R. and D. Pauly, eds.), licensed CC BY-NC 4.0.Last updated 27 September 2026