(total): 13 - 14;
soft rays: 19 - 23. Gill membranes not forming fold across isthmus. Tentacles on nasal openings and above eyes. No canine teeth in upper jaw. Dentaries joined by ligament. Frontals not covered by parietals. Spawning males chocolate-brown, with bulb glands on spiny anal fin rays. Lateral line continuous with branches but discontinuous posteriorly.