Inhabit fast rocky riffles of creeks and small to medium rivers (Ref. 33799); also found in medium to large streams (Ref. 10294). Adults occur in the deepest portions of riffles with moderately strong current. Prey mainly on insects (midge and other dipteran larvae, mayfly nymphs, and smaller quantities of water mites and caddisfly larvae (Ref. 10294)). Feed by pecking small prey from the surface of rocks (Ref. 33799).