Inhabit rocky riffles of creeks and small to medium rivers (Ref. 5723, 10294); also found in small to medium upland streams (Ref. 10294). Adults feed on midge, blackfly, and caddisfly larvae, mayfly nymphs, isopods, and amphipods (Ref. 10294). Eggs are found clustered on underside of stone and guarded by males (Ref. 7043).